crease
in thickness from the hinge to the edge, and the inside lip of the
shell shows the gradual union of the three superimposed layers. The two
outer layers are formed by the thick edge of the mantle, the remaining
portion—or nearly the entire surface—of this organ secretes the nacrai
layer.
Not
only is the interior of the shell made lustrous and beautiful, but this
tendency is exerted toward all objects that come in contact with the
soft body of the mollusk, either by intrusion simply within the shell,
or deeply within the organs and tissues of the animal itself. All
foreign bodies—such as small parasites, diatoms, minute pebbles, etc.,
—irritate the tender tissues of the mollusk, and stimulate the pearly
formation which in course of time covers them. At first the nacreous
covering is very thin ; but with added layer after layer the thickness
is enhanced, and the size of the object increases as long as it remains
undisturbed and the mollusk is in healthful growth.
Chemically
considered, aside from the nucleus, the structure of pearls is
identical in composition with that of the nacre of the shell in which
they are formed. Analyses have shown that those from the fresh-water
mussels of England and Scotland, and from the pearl-oysters of
Australia and of Ceylon, have nearly identical composition in the
proportion of about 5.94 per cent, of organic matter, 2.34 of water,
and 91.72 per cent, of carbonate of lime.1 The specific
gravity ranges from nearly 2 to about 2.75, increasing with the deposit
of the nacreous coatings. The following summary by Von Hessling2 shows the results of certain determinations of specific gravity :